Last update: 03 December 2016


Custom Search

ABCDEFGHIJKLMNOPQRSTU – V – W – X – YZ




  • Abdoli L., Kamrani E., Abdoli A. and Kiabi B., 2009 – Length-weight relationships for three species of mudskippers (Gobiidae: Oxudercinae) in the coastal areas of the Persian Gulf, Iran – Journal of Applied Ichthyology, 25(2): 236–237.

  • Afzelius B.A., 1956 – Seitenorgane und Schleimkalanknocken bei Periophthalmus koelreuteri und Gobius minutusZeitschrift für Anatomie und Entwicklungsgeschichte, 119: 470-484.

  • Agorreta A., Rüber L., 2012 – A standardized reanalysis of molecular phylogenetic hypotheses of Gobioidei – Systematics and Biodiversity, 10: 375-390.

  • Agorreta A., San Mauro D., Schliewen U., Van Tassell J.L., Kovacic M., Zardoya R., Rüber L., 2013 – Molecular phylogenetics of Gobioidei and phylogenetic placement of European gobies – Molecular Phylogenetics and Evolution, 69(3): 619-633.

  • Aguilar N.M., 2000 – Comparative physiology of air–breathing gobies unpubl. Ph D. dissertation, Univ. of California, San Diego, United States of America, 201 pp.

  • Akihito, Iwata A., Kobayashi T., Ikeo K., Imanishi T., Ono H., Umehara Y., Hamamatsu C., Sugiyama K., Ikeda Y., Sakamoto K., Fumihito A., Ohno S., Gojobori T., 2000 – Evolutionary aspects of gobioid fishes based upon a phylogenetic analysis of mitochondrial cytochrome b genes – Gene (Elsevier), 259: 5–15.

  • Al-Kadhomiy, N. K. and Hughes, G. M. 1988 – Histological study of different regions of the skin and gills in the mudskipper, Boleophthalmus boddarti with respect to their respiratory function – Journal of the Marine Biological Association of the United Kingdom, 68: 413–422.

  • Allen G. (ed), 1991 – Field Guide to the Freshwater Fishes of New Guinea – Christensen Research Institute.

  • Allen G. (ed), 1997 – Marine Fishes of the Great Barrier Reef and South–East Asia – Western Australian Museum.

  • Allen, G.R., S.H. Midgley and M. Allen (eds), 2002 – Field Guide to the Freshwater Fishes of Australia – Western Australian Museum, Perth, Western Australia. 394 p.

  • Arens N.C., Strömberg C. and Thompson A., 1998 – Lab IV, Early land plants – World Wide Web electronic publication, www.ucmp.berkeley.edu/IB181/VPL/Elp/ElpTitle.html, visited (11/2005).

  • Axelrod H.R., Axelrod G.S., Burgess W.E., Scott B.M., Pronek N. and Walls J.G., 2004 – Dr. Axelrod's Atlas of Freshwater Aquarium Fishes – 10th edition, T.F.H. Publ. Inc.




  • Baeck G.W., Takita T. and Yoon Y.H., 2008 – Lifestyle of Korean mudskipper Periophthalmus magnuspinnatus with reference to a congeneric species Periophthalmus modestusIchthyological Research, 55: 43–52.

  • Bajpai S. and Kapur V.V., 2004 – Oldest known gobiids from Vastan Lignite Mine (early Eocene), Surat District, Gujarat – Current Science, 87(4): 433–435.

  • Barak, N.A.–E., Salman, N.A. and Ahmad, S.M.,1994 – The piscivorous feeding of mudskipper Periophthalmus waltoni Koumans from Khor Al–Zubair, northwest Arabian Gulf – Pakistan Journal of Zoology, 26(3): 280–283.

  • Bauchot, M.-L., Whitehead P.J.P. and Monod T., 1982 – Date of publication and authorship of the fish names in Eydoux & Souleyet's zoology of La Bonite, 1841-1852 – Cybium 3e série. Bulletin de la Société Française d'Ichtyologie, 6(3): 59–73.

  • Bauchot, M.-L., M. Desoutter, D.F. Hoese and H.K. Larson, 1991 – Catalogue critique des types de Poissons du Muséum national d'Histoire naturelle. (Suite) Sous–ordre des Gobioidei – Bull. Mus. Natl. Hist. Nat. Ser. 4, Sect. A v. 13 (no. 1–2, suppl.): 1–82.

  • Berg L.S., 1949 – Presnovodnye ryby Irana i sopredel'nykh stran [Freshwater fishes of Iran and adjacent countries] – Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, 8: 783–858.

  • Berti R., Chelazzi L., Colombini I. and Ercolini A.,1992 – Direction–finding ability in a mudskipper from the delta of the Tana river (Kenya) – Tropical Zoology, 5: 219–228.

  • Berti R., Colombini I, Chelazzi L. and Ercolini A., 1994 – Directional orientation in Kenyan population of Periophthalmus sobrinus Eggert: experimental analysis of the operating mechanism – Journal of Experimental Marine Biology and Ecology, 181: 135–141.

  • Betancur-R R., Wiley E.O., Miya M., Lecointre G., Bailly N., Ortí G., 2014 New and revised classification of bony fishes. Version 2. (Electronic version accessed 24.06.14).

  • Bhatt N.Y., Patel S.J., Patel D.A. & Patel H.P., 2009 – Burrowing Activities of goby fish in the recent intertidal mud flats along the Navinal Coast, Kachchh, Western India – Journal Geological Society of India, 74: 515-530.

  • Bianchi G., 1986 – Fichas FAO de identifacao de espécies para propósitos comerciais. Guia de campo para as espécies comerciais marinhas e de águas salobras de Angola– Preparado com o apoio da NORAD e da FAO (FIRM) Programa Regular, FAO, Rome. 184 p.

  • Bianconi G.G., 1855–59 – Specimina zoologica mosambicana quibus vel novae vel minus notae animalium species illustrantur – Bononiae, 1850–1870 [Fascicles X (1855), XII (1857), XIII (1859) involve fishes]. Mosambique Zoology: 1–363, 48 col pls.

  • Bleeker P., 1849 – Bijdrage tot de kennis der Blennioïden en Gobioïden van der Soenda–Molukschen Archipel, met beschrijving van 42 nieuwe soorten – Verhandelingen van het Bataviaasch Genootschap Kunst. Wet., 22(6): 1–40.

  • Bleeker P., 1851a – Derde bijdrage tot de kennis der ichthyologische fauna van Borneo, met beschrijving van eenige nieuwe soorten van zoetwatervisschen – Natuurkdig Tijdschrift voor Nederlandsch Indië, 2: 57–70.

  • Bleeker P., 1851b – Bijdrage tot de kennis der ichthyologische fauna van Borneo, met beschrijving van 16 nieuwe soorten van zoetwatervisschen – Natuurkdig Tijdschrift voor Nederlandsch Indië, 1: 1–16.

  • Bleeker P., 1852 – Nieuwe bijdrage tot de kennis der ichthyologische fauna van het eiland Banka – Natuurkdig Tijdschrift voor Nederlandsch Indië, 3: 715–738.

  • Bleeker P., 1853 – Vierde bijdrage tot de kennis der ichthyologische fauna van Amboina – Natuurkundig Tijdschrift voor Nederlandsch Indië, 5: 317–352.

  • Bleeker P., 1854 – Ichthyologische waarnemingen, gedaan op verschillende reizen in de residentie Banten – Natuurkundig Tijdschrift voor Nederlandsch Indië, 7: 309–326.

  • Bleeker P., 1874 – Esquisse d'un systeme naturel des Gobioides – Archives Nerlandaises des Sciences Exactes et Naturelles, 9: 289–331.

  • Bleeker P., 1983 – Atlas Ichthyologique des Indes Orientales Néêrlandaises. Plates originally prepared for planned tomes XI-XIV. Smithsonian Institution Press, Washington, D.C.

  • Bloch M.E. and Schneider J.G., 1801 – M.E. Blochii, Systema Ichthyologiae iconibus CX illustratum. Post obitum auctoris opus inchoatum absolvit, correxit, interpolavit Jo. Gottlob Schneider, Saxo. Berolini. Sumtibus Auctoris Impressum et Bibliopolio Sanderiano Commissum – i–lx + 1–584.

  • Blyth E., 1858 – Report of Curator, Zoological Department, for May, 1858 – J. Asiatic Soc. Bengal, 27(3): 267–290. [Fishes are discussed on pp. 270–272 and 281–290.]

  • Blyth E., 1860 – Report on some fishes received chiefly from the Sitang River and its tributary streams, Tenasserim Provinces – J. Asiatic Soc. Bengal: 138–174.

  • Bostock J. & Riley H.T., 1855 – The Natural History, Pliny the Elder. Taylor and Francis, London.

  • Brillet C., 1969a – Etude du comportement constructeur des poissons amphibies Periophthalmidae – Terre et la Vie, 23 (4): 496–520.

  • Brillet C., 1969b – Comportement territoriale et hierarchie sociale chez le poisson amphibie du genre Periophthalmus (Téléostéen, Gobioidea) – Compt. Rend. Hebd. des Séan. de l' Acad. des Sc., Ser. D, 269: 1322–1325.

  • Brillet C., 1970 – Relations entre comportement sexuel, territoire et aggressivitè chez le periophtalmes – Compt. Rend. Hebd. des Séan. de l' Acad. des Sc., Ser. D, 270: 1507–1510.

  • Brillet C., 1975 – Relations entre territoire et comportement agressif chez Periophthalmus sobrinus Eggert (Pisces, Periophthalmidae) au laboratoire et en milieu naturel – Zeitschr. Tierpsychol., 39: 283–331.

  • Brillet C., 1976 – Structure du terrier, reproduction et comportement des jeunes chez le poisson amphibie Periophthalmus sobrinus Eggert – Terre et la Vie, 30 (3): 465–483.

  • Brillet C., 1980a – Agonistic behaviour of an amphibious fish, Periophthalmus sobrinus Eggert. Frequency of the various elements of the repertoire – Biology of Behaviour/Biologie Comport., 5 (4): 297–315.

  • Brillet C., 1980b – Comportement sexuel du poisson amphibie Periophthalmus sobrinus Eggert: ses rapports avec le comportement agonistique – Revue d'Ecologie, Terre et la Vie, 34 (3): 427–468.

  • Brillet C., 1981 – Comportement agonistique du poisson amphibie Periophthalmus sobrinus Eggert. Analyse quantitative 2. Influence de la taille et du sex sur l'establissement des relations de dominance/subordination et sur l'utilisation du repertoire agonistique – Biology of Behaviour/Biologie Comport., 6 (1): 35–57.

  • Brillet C., 1983 – Comportement agonistique du poisson amphibie Periophthalmus sobrinus Eggert. Analyse quantitative 3. Role de nageoires dorsales dans l'establissement des relations de dominance–subordination – Biology of Behaviour/Biologie Comport., 8 (1): 49–66.

  • Brillet C., 1984a – Etude comparative de la parade nuptiale chez deux especes de Poissons amphibies sympatriques (Pisces, Periophthalmidae) – C. R. Hebd. Séances Acad. Sc. Serie D, Paris, 198 (12): 347–350.

  • Brillet C., 1984b – Habitat et comportement du poisson amphibie Periophthalmus koelreuteri africanus Eggert. A Tulear comparison avec le Periophthalmus sobrinus Eggert sympatrique – Terre et la Vie, 39 (3): 337–346.

  • Bucholtz R.H. and Meilvang A.S., 2005 – Biology and utilization of the mudskipper Pseudapocryptes elongatus in the Mekong River Delta, with aspects on other mudskippers in the area – UM.Sc. Hons. Thesis, Department of Marine Ecology and Centre for Tropical Ecosystem Research, University of Aarhus, Denmark, 99 pp.

  • Bucholtz R.H., Meilvang A.S., Cedhagen T. & Christensen J.T., 2009 – Biological observations on the mudskipper Pseudapocryptes elongatus in the Mekong Delta, Vietnam – Journal of the World Aquaculture Society, 40(6): 711-723.

  • Bullini L., Pignatti S. and Virzo De Santo, 1998 – Ecologia generale – UTET, Torino, 519 pp.




  • Cantor T., 1842 – General features of Chusan, with remarks on the flora and fauna of that island – The Annals and Magazine of Natural History, 9: 481–493.

  • Cantor T., 1850 – Catalog of Malaysan Fishes – Journal of the Asiatic Society of Bengal , 18(2): 983–1443.

  • Carroll R.L., Irwin J. and Green D.M., 2005 – Thermal physiology and the origin of terrestriality in vertebrates – Zoological Journal of the Linnean Society, 143: 345–358.

  • Castelnau F.L., 1873 – Contribution to the ichthyology of Australia. Nos. III thru IX [with subtitles, indexed as one work] – Proc. Zool. Acclim. Soc. Victoria: 37–158.

  • Castelnau F.L., 1876 – Researches on the fishes of Australia. pp. 1–52 – In: Official Record, Philadelphia Centennal Exhibition of 1876, Intercolonial Exhibition essays 1875–1876, Vol. 2.

  • Chakrabarty P., Davis M.P. and Sparks J.S. (2012) – The first record of a trans-oceanic sister-group relationship between obligate vertebrate troglobites – PLOS ONE, 7(8): e44083. doi:10.1371/journal.pone.0044083.

  • Chen H., Polgar G., Yin W. and Fu C.-Z. (2014) – Cryptic species and evolutionary history of Boleophthalmus pectinirostris complex along the northwestern Pacific coast – Acta Hydrobiologica Sinica, 38(1): 75-86 (in Cinese con sommario inglese).

  • Chen Y.D. and Ting Y.Y., 1986 – The Preliminary report on inducing breeding experiments with mudskipper Boleophthalmus pectinirostris L., by injection of HCG – Bulletin of the Taiwan Fish Res. Inst., 40: 53–59 (in Cinese con sommario inglese).

  • Chen S. Hong W., Zhang Q. and Su Y., 2007 – Why does the mudskipper Boleophthalmus pectinirostris form territories in farming ponds? – Journal of the Marine Biological Association of the United Kingdom, 87: 615–619.

  • Chew S.F. and Ip Y. K., 1992 – Cyanide detoxification in the mudskipper, Boleophthalmus boddaertiJournal of Experimental Zoology, 261: 1–8.

  • Chew S.F., Goh E., Lim C.E. and Ip Y. K., 1998 – Cyanide exposure affects the production and excretion of ammonia by the mudskipper Boleophthalmus boddaertiComparative Biochemistry and Physiology Part C, 120: 441–448.

  • Chew S.F., Hong L.N., Wilson J.M., Randall D.J. and Ip Y. K., 2003 – Alkaline Environmental pH has no effect on ammonia excretion in the mudskipper Periophthalmodon schlosseri but inhibits ammonia excretion in the related species Boleophthalmus boddaertiPhysiological and Biochemical Zoology, 76(2): 204–214.

  • Chung, E.–Y.; An, C.–M. and Lee, T.–Y., 1991 – Sexual maturation of the bluespotted mud hopper, Boleophthalmus pectinirostris (Linnaeus) – Bulletin of the Korean Fisheries Society, 24(3): 167–176. (In Korean with English summary)

  • Clack J.A., 2002 – Gaining Ground: The Origin and Evolution of Tetrapods – Farlow J.O. (ed), Indiana University Press, Bloomington and Indianapolis: 369 pp.

  • Clayton D.A., 1985 – Ecology of mudflats with particular reference to those on the northern Arabian Gulf – In: Marine environment and pollution. Proceedings of the Conference on the Marine Environment and Pollution; R. Halwagy, D. Clayton, and M. Behbehani (eds.). Kuwait University, Kuwait Foundation for the Advancement of Science and Environmental Protection Council, Kuwait City, Kuwait, p. 83–96.

  • Clayton D.A., 1987 – Why mudskippers build walls – Behaviour, 102: 185–195.

  • Clayton D.A., 1993 – Mudskippers – Oceanogr. Mar. Bull. Annu. Rev. A.D. Ansell, R.N. Gibson and Margaret Barnes (eds), UCL Press, 31: 507–577.

  • Clayton D.A. and Snowden R., 2000 – Surface activity in the mudskipper, Periophthalmus waltoni in relation to prey activity and environmental factors – Tropical Zoology, 13(2): 239–249.

  • Clayton D.A. and Vaughan T.C., 1982 – Pentagonal Territories of the Mudskipper Boleophthalmus boddaerti (Pisces, Gobiidae) – Copeia, 1: 232–234.

  • Clayton D.A. and Vaughan T.C., 1986 – Territorial acquisition in the mudskipper Boleophthalmus boddarti (Pisces: Gobiidae) on the mudflats of Kuwait – Journ. Zool. (A), London, 209: 501–519.

  • Clayton D.A. and Vaughan T.C., 1988 – Ethogram of Boleophthalmus boddarti (Pallas), a mudskipper found on the mudflats of Kuwait – Journal of the University of Kuwait (Sciences), 15 (1): 115–138.

  • Clayton D.A. and Wright J.M., 1989 – Mud–walled territories and feeding behaviour of Boleophthalmus boddarti (Pisces, Gobiidae), on the mud flats of Kuwait – Journal Ethol., 7 (2): 91–95.

  • Colombini I; Berti R.; Nocita A. and Chelazzi I., 1995 – Environmental factors influencing the zonation and activity patterns of a population of Periophthalmus sobrinus Eggert in a Kenian mangrove – Journal of Experimental Marine Biology and Ecology, 190: 135–149.

  • Colombini I; Berti R.; Nocita A. and Chelazzi I., 1996 – Foraging Strategy of the Mudskipper Periophthalmus sobrinus Eggert in a Kenian mangrove – Journal of Experimental Marine Biology and Ecology, 197 (2): 219–235.

  • Costa D.P., 1999 – The role of physiology in the behavior of diving mammals: insights from animals in nature – EUBS 99 Proceedings 25th Annual Meeting Haifa and Eilat Israel: 233–239.

  • Cuvier G., 1816 – Le Règne Animal distribué d'après son organisation pour servir de base à l'histoire naturelle des animaux et d'introduction à l'anatomie comparée. Les reptiles, les poissons, les mollusques et les annélides – Edition 1. i–xviii + 1–532.

  • Cuvier G. and Valenciennes A., 1837 – Histoire naturelle des poissons. Tome douzième. Suite du livre quatorzième. Gobioïdes. Livre quinzième. Acanthoptérygiens à pectorales pédiculées – i–xxiv + 1–507 + 1 p.

  • Czerkas S.J. and Crerkas S.A., 1991 – Dinosaurs: A Global View – BDD Promotional Books Company.




  • Darumas U. and Tantichodok P., 2002 – A new species of Mudskipper (Gobiidae: Oxudercinae) from southern Thailand – Phuket Marine Biological Center Research Bulletin, 64: 101–107.

  • Day F., 1867 – On some new or imperfectly known fishes of Madras – Proceedings of the Zoological Society of London – 1867: 935–942.

  • Day F., 1871 – On the fishes of the Andaman Islands – Proceedings of the General Meetings for Scientific Business of the Zoological Society London; 1870: 677–705.

  • Day F., 1876 – The fishes of India; being a natural history of the fishes known to inhabit the seas and fresh waters of India, Burma, and Ceylon – Part 2: 169–368 + pls. 41–78.

  • Day F., 1889 – The fauna of British India, including Ceylon and Burma – Fishes, Vol. 2. Taylor & Francis, London.

  • Davis W.P., Taylor D.S. & Turner B.J., 1990 – Field observations of the ecology and habits of mangrove Rivulus (Rivulus marmoratus) in Belize and Florida – Ichthyological Exploration of Freshwaters, 1: 123–134.

  • De Vis C.W., 1884 – Fishes from South Sea islands – Proc. Linn. Soc. N.S.W.: 445–457.

  • Diesselhorst G., 1938 – Horversuche an Fischen ohne Weberschen Apparat – Zeitschrift für Vergleichende Physiologie, 25: 748–783.

  • Diesselhorst G., 1938 – Horversuche an Fischen ohne Weberschen Apparat – Zeitschrift für Vergleichende Physiologie, 25: 748–783.

  • Dinh T.D., Ambak M.A., Hassan A. & Phuong N.T., 2007 – Biology and population dynamics of the goby Pseudapocryptes elongatus in the coastal mud flat areas of the Mekong Delta, Vietnam – Pakistan Journal of Biological Sciences, 10(19): 3284–3294.

  • Dotsu Y., 1961 – The bionomics and life history of the gobioid fish, Apocryptodon bleekeri (Day) – Bulletin of the Faculty of Fisheries, Nagasaki University (10): 127–131.

  • Duméril A.H.A., 1861 – Poissons de la côte occidentale d'Afrique – Archives du Museum National d'Histoire Naturelle (Paris) v. 10 [for 1858]: 241–268, Pls. 20–23. [according to the Catalog of Fishes by Eschmeyer, apparently published in 1861].




  • Eccles D.H., 1992 – FAO species identification sheets for fishery purposes. Field guide to the freshwater fishes of Tanzania – Prepared and published with the support of the United Nations Development Programme (project URT/87/016). FAO, Rome. 145 p.

  • Egborge A.B.M., 1993 – Biodiversity of aquatic fauna of Nigeria – Natural Resources Conservation Council (Nigeria), 173 p.

  • Eggert B., 1929 – Bestimmungstabelle und Beschreibung der Arten der Familie Periophthalmus – Zeitschrift für Wissenschaftliche Zoologie, 133(1,2): 398–410, Pls. 8–9.

  • Eggert B., 1935 – Beitrag zur Systematik, Biologie und geographischen Verbreitung der Periophthalminae. Ergebnisse einer durch die Notgemeinschaft der Deutschen Wissenschaft ermöglichten Reise nach Niederländisch-Indien 1926-1927 und der Sundaexpedition der Notgemeinschaft der Deutschen Wissenschaft 1929-1930 – Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere (Jena): 29–116.

  • Eiseley, 1957 – The Immense Journey – Random House, New York, 210 pp.

  • Ellison A., Farnsworth E. and Merkt R.E., 1999 – Origin of Mangrove Ecosystems and the mangrove biodiversity anomaly – Global Ecology and Biogeography, 8: 95–115.

  • Eschmeyer W.N., 1998 (ed) – Catalog of Fishes – Center for Biodiversity Research and Information, Spec. Publ. 1. California Academy of Sciences. vols. 1–3: 1–2905

  • Etim L., Brey T. and Arntz W., 1996 – A seminal study of the dynamics of a mudskipper (Periophthalmus papilio) population in the Cross river, Nigeria – Netherlands Journal of Aquatic Ecology 30(1): 79–96.

  • Etim L., King R.P. and Udo M.T., 2002 – Breeding, growth, mortality and yield of the mudskipper Periophthalmus barbarus (Linnaeus 1766) (Teleostei: Gobiidae) in the Imo River estuary, Nigeria – Fisheries Research 56: 227–238.

  • Evans D.H., Claiborne J.B. and Claiborne G.K., 1999 – Osmoregulation, acid–base regulation, and nitrogen excretion – In: Horn M.H., Martin K.L.M. & Chotkowski M.A. (eds) – Intertidal fishes: life in two worlds – Academic Press, San Diego California, 399 pp: 79–96.

  • Eydoux J.F.T. and Souleyet F.L.A., 1842/1848 – Poissons. Pp. 155–216. In: Voyage autour du monde exécuté pendant les années 1836 et 1837 sur la corvette La Bonite, commandée par M. Vaillant Zoologie, Vol. 1 (part 2). Paris. Voyage Bonite: i–iv, i–xxxix, 1–334, Pls. 1–10.




  • Ferraris C.J. Jr., 1995 – On the validity of the name Pseudapocryptes lanceolatus (Bloch and Schneider, 1801) (Osteichthyes: Percomorpha: Gobiidae) – Copeia, 1995(4): 984.

  • Fineman–Kalio A.S and Alfred–Okciya J.F., 1989 – Towards large scale farming of the mudskipper in the brackish environment – In: Proceeding of 4th annual conference of the Nigerian association for aquatic sciences at the Dept. of wildlife and fisheries management, Univ. of Ibadan, Nigeria: 7 p.

  • Fowler H.W., 1934 – Zoological results of the third De Schauensee Siamese Expedition, Part I.––Fishes – Proc. Acad. Nat. Sci. Phila.: 67–163.

  • Fowler H.W., 1935 – Zoological results of the third De Schauensee Siamese Expedition, Part VI.––Fishes obtained in 1934 – Proc. Acad. Nat. Sci. Phila.: 89–163.

  • Fowler H.W., 1937 – Zoological results of the third De Schauensee Siamese Expedition, Part VIII,––Fishes obtained in 1936 – Proc. Acad. Nat. Sci. Phila.: 125–264.

  • Froese R. and Pauly D. (eds), 2005 – Fishbase – World Wide Web electronic publication, www.fishbase.org, visited (11/2005).




  • Ghanbarifardi M., Aliabadian M., Esmaeili H.R. and Polgar G., 2014 – Morphological divergence in the Walton’s mudskipper, Periophthalmus waltoni Koumans, 1941, from the Persian Gulf and Gulf of Oman (Gobioidei: Gobiidae) – Zoology in the Middle East, 60(2): 133-143.

  • Garey W.F., 1962 – Cardiac responses of fishes in asphyxic environments – The Biological Bulletin, 122: 362–368.

  • Gee J.H. and Gee P.A., 1995 – Aquatic surface respiration, buoyancy control and the evolution of air breathing in gobies (Gobiidae: Pisces) – Journal of Experimental Biology, 198: 79–89.

  • Ghosh R.S., Ebbs S.D., Bushey J.T., Neuhauser E.F. and Wong-Chong G.M., 2006 – Cyanide in nature – In: Dzombak D.A., Ghosh R.S. and Wong-Chong G.M. (eds) Cyanide in water and soil: chemistry, risk and management, CRC Press, Taylor & Francis Group, Boca Raton, pp. 225–236.

  • Gmelin J.F., 1789 – Caroli a Linné, Systema Naturae – Tom. I, pars III: 1196–1206. Lipsiae (Beer)

  • Gordon M.S., Boetius J., Evans D.H., Mc Carthy R. & Oglesby L.C., 1969 – Aspects of the physiology of terrestrial life in amphibious fishes I. The mudskipper Periophthalmus sobrinusThe Journal of Experimental Zoology, 50: 141–149.

  • Gordon M.S.; Ng W.W.S. & Yp A.K.W., 1978 – Aspects of the physiology of terrestrial life in amphibious fishes III. The Chinese mudskipper Periophthalmus cantonensisJournal of Experimental Biology, 72: 57–75.

  • Graham J.B., 1971 – Aerial vision in amphibious fishes – Fauna (Rancho Mirage, California), 3: 14–23.

  • Graham J.B., 1997 (ed) – Air–breathing Fishes. Evolution, Diversity and Adaptation – Academic Press, San Diego California, 299 pp.

  • Graham J.B., Aguilar N., Dudley R. and Gans C., 1997 – The Late Paleozoic atmosphere and the ecological and evolutionary physiology of tetrapods – In: Sumida S. & Martin K.L.M. (eds) – Amniote Origins: Completing the Transition to Land – Academic Press, San Diego California: 141–167.

  • Graham J.B. and Lee H.J., 2004 – Breathing air in air: in what ways might extant amphibious fish biology relate to prevailing concepts about early tetrapods, the evolution of vertebrate air breathing, and the vertebrate land transition? – Physiological and Biochemical Zoology, 77(5): 720–731.

  • Graham J.B., Lee H.J. and Wegner N.C., 2007 – Transition from water to land in an extant group of fishes: air breathing and the acquisition sequence of adaptations for amphibious life in oxudercine gobies – In: Fernandes M.N., Rantin F.T., Glass M.L., & Kapoor B.G. (eds) Fish Respiration and Environment. Science Publishers, Enfield, pp 255–288.

  • Gregory R.B., 1977 – Synthesis and total excretion of waste Nitrogen by fish of the Periophthalmus (mudskipper) and Scartelaos families – Comparative Biochemistry and Physiology, 57A: 33–36.

  • Grizzle J.M. & Thiyagarajah A., 1987 – Skin histology of Rivulus ocellatus marmoratus, apparent adaptation for aerial respiration – Copeia : 237–240.

  • Günther A., 1861 – Catalogue of the fishes in the collection of the British Museum. Vol. 3, London.

  • Günther A., 1867 – Description of a new species of ApocryptesThe Annals and Magazine of Natural History, 20: 117.




  • Hamilton F. [Buchanan, or Hamilton-Buchanan, or Buchanan-Hamilton], 1822 – An account of the fishes found in the river Ganges and its branches – Edinburgh & London. i–vii + 1–405.

  • Harms J.W., 1929 – Die Realization von Genen und die consecutive Adaptation. 1: Phasen in der Differenzierung der Anlagenkomplexe und die Frage der Landtierwerdung – Zeitschrift für wissenschaftliche Zoologie, 133: 211–397.

  • Harris V.A., 1961 – On the locomotion of the mudskipper Periophthalmus koelreuteri (Pallas): Gobiidae – Proceedings of the Zoological Society of London, 134: 107–135.

  • Harrison I.J. and Miller P.J., 1992– Gobiidae. p. 798–821. In: Levêque C., Paugy D. and Teugels G.G. (eds.) Faune des poissons d'eaux douces et saumâtres d'Afrique de l'Ouest, Tome 2. Coll. Faune Tropicale n° 28. Musée Royal de l'Afrique Centrale, Tervuren, Belgique and O.R.S.T.O.M., Paris, France, 902.

  • Harzauser M., Kroh A., Mandic O., Piller W.E., Glich U., Reuter M., Berning B., 2007 – Biogeographic responses to geodynamics: a key study all around the Oligo-Miocene Tethyan seaway – Zoologischer Anzeiger, 246: 241–256.

  • Henderson D. (ed), 1999 – Earth History Illustrations – World Wide Web electronic publication, http://gallery.in–tch.com/%7Eearthhistory/index.html, visited (11/2005).

  • Hase A., 1914 – Die Fische der Deutschen Grenzexpedition 1910 in das Kaiser–Wilhelms– Land, Neu Guinea – Jena Zeitschr. Naturw.: 525–548.

  • Herre A.W.C.T., 1927 – Gobies of the Philippines and China Sea – Monogr. Bur. Sci. Manila, 23: 352 pp.

  • Herre A.W.C.T., 1932 – Fishes from Kwangtung Province and Hainan Island, China – Lingnan Science Journal Canton, 11: 423–443

  • Hoda S.M.S, 1986 – Maturation and fecundity of the mudskipper Boleophthalmus dussumieri Cuv. & Val. from the Karachi coast – Mahasagar, Bulletin of the National Institute of Oceanography, 19(1): 73–78.

  • Hoda S.M.S, 1987 – Relative growths of body parts and length-weight relationships in Boleophthalmus dussumieri and B. dentatus of Karachi coast – Indian Journal of Fisheries, 34(1): 120–127.

  • Hoda S.M.S. and Akhtar Y., 1985 – Maturation and fecundity of the mudskipper Boleophthalmus dentatus in the northern Arabian Sea. Indian Journal of Fisheries, 32: 64–74.

  • Hoese D.F., 1984 – Gobioidei: relationships – In: Moser H.G., Richards W.J., Cohen D.M., Fahay M.P., Kendall A.W. Jr. and Richardson S.L. (eds) – Ontogeny and systematics of fishes – Spec. Publ. No. 1, Amer. Soc. Ichthy. and Herpet., Allen Press, Lawrence Kansas, 760 pp.

  • Hogarth P.J. (ed), 1999 – The Biology of Mangroves – Oxford Univ. Press, Oxford, 228 pp.

  • Holly M., 1929 – Drei neue Fischformen aus Persien – Anzeiger der Akademie der Wissenschaften in Wien v. 66 (7): 62–64.

  • Hong W., Dai Q.–Y., Zhang Q.–Y., Zhang J. and Cai Y.–Y., 1988 – Observations on the early development of the mudskipper Boleophthalmus pectinirostris (Linnaeus) – Tropical Oceanology / Redai Haiyang, 2: 1–8. (In Chinese with English summary)

  • Hong, W. and Wang Y.–L., 1989 – Induction of ovulation in female mudskipper (Boleophthalmus pectinirostris), by HCG, analogue of LHRH and Pimozide – Chinese Journal of Oceanology and Limnology, 7(3): 283–287. (In Chinese with English summary)

  • Hora S.L., 1933 – Respiration in fishes – Journal of the Bombay Natural History Society, 36: 538–560.

  • Hora S.L., 1934 – The systematic position of Hamilton's species of gobioid fishes from the Ganges – Records of the Indian Museum, Calcutta, 36: 483–490.

  • Hora S.L., 1935 – Ecology and bionomics of the gobioid fishes of the Gangetic Delta – Comptes Rendus, XII Congrès International de Zoologie, Vol. II, Lisbon 1935: 841–865.

  • Horn M.H., Martin K.L.M. and Chotkowski M.A. (eds), 1999 – Intertidal Fishes: Life in Two Worlds – Academic Press, San Diego, 399 pp.




  • Ikebe,Y. and Oishi T., 1996 – Correlation between environmental parameters and behaviour during high tides in Periophthalmus modestusJournal of Fish Biology, 49(1): 139–147.

  • Inger R.F., 1957 – Ecological aspects of the origin of tetrapods – Evolution, 11: 373–376.

  • Ip Y.K., Low W.P., Lim A.L.L. and Chew S.F., 1990 – Changes in Lactate Content in the Gills of the Mudskipper Periophthalmus chrysospilos and Boleophthalmus boddaerti in Response to Environmental Hypoxia – Journal of Fish Biology, 36(4): 481–487.

  • Ip Y.K. and Low W.P., 1990 – Lactate Production in the Gills of Mudskipper Periophthalmodon schlosseri Exposed to Hypoxia – Journal of Experimental Zoology, 253(1): 99–101.

  • Ip Y.K., Lim C.B., Chew S.F., Wilson J.M. and Randall D.J., 2001 – Partial amino acid catabolism leading to the formation of Alanine in Periophthalmodon schlosseri (mudskipper): a strategy that facilitates the use of amino acids as an energy resource during locomotory activity on land – Journal of Experimental Biology, 204: 1615–1624.

  • Ip Y.K., Chew S.F. and Randall, D.J., 2004a – Five tropical air–breathing fishes, six different strategies to defend against ammonia toxicity on land – Physiological and Biochemical Zoology, 77(5): 768–782.

  • Ip Y.K., Randall, D.J., Kok T.K.T., Barzaghi C., Wright P.A., Ballantyne J.S., Wilson J.M. and Chew S.F., 2004b – The giant mudskipper Periophthalmodon schlosseri facilitates active NH4+ excretion by increasing acid excretion and decreasing NH3 permeability in the skin – The Journal of Experimental Biology, 207: 787–801.

  • Ip Y.K., Kuah S.S.L. and Chew S.F., 2004c – Strategies adopted by the mudskipper Boleophthalmus boddaerti to survive exposure in normoxia or hypoxia – Physiological and Biochemical Zoology, 77(5): 824–837.

  • Ishimatsu A., Hishida Y., Takita T., Kanda T., Oikawa S., Takeda T. and Khoo K.H., 1998 – Mudskipper Store Air in Their Burrows – Nature, 391: 237–238.

  • Ishimatsu A., Aguilar N.M., Ogawa K., Hishida Y., Takeda T., Oikawa S., Kanda T. and Huat K.K., 1999 – Arterial blood gas levels and cardiovascular function during varying environmental conditions in a mudskipper, Periophthalmodon schlosseriJournal of Experimental Biology, 202: 1753–1762.

  • Ishimatsu A., Takeda T., Kanda T., Oikawa S. and Khoo K.H., 2000 – Burrow environment of mudskippers in Malaysia – Journal of Bioscience, 11 (1,2): 17–28.

  • Ishimatsu A., Yoshida Y., Itoki N., Takeda T., Lee H.J. and Graham J.B., 2007 – Mudskippers brood their eggs in air but submerge them for hatching – The Journal of Experimental Biology, 210: 3946–3954.

  • Islam Md.S., Hossain Md.S., Chowdhury S.R. and Khan Y.S.A., 2006 – Effect of sublethal doses of DDT on the liver of Apocryptes bato (Bleeker, 1874) – Chiang Mai Journal of Science, 33(3): 211–215.

  • Itoki N., Sakamoto T., Hayashi M., Takeda T. and Ishimatsu A., 2012 – Morphological responses of mitochondria-Rich cells to hypersaline environment in the Australian mudskipper, Periophthalmus minutusZoological Science, 29(7): 444–449.

  • Iwata K., 1988 – Nitrogen metabolism in the mudskipper Periophthalmus cantonensis. Changes in free aminoacids and related compounds in various tissues, under conditions of ammonia loading with special reference to its high ammonia tolerance – Comparative Biochemistry and Physiology (A): Comparative Physiology, 91(33): 499–508.

  • Iwata K., Kakuta I., Ikeda M., Kimoto S. and Wada N., 1981 – Nitrogen metabolism in the mudskipper Periophthalmus cantonensis: a role of free aminoacids in detoxication of ammonia produced during its terrestrial life – Comparative Biochemistry and Physiology (A): Comparative Physiology, 68(4): 589–596.




  • Jaafar Z., and Parenti L.R., 2016 – Systematics of the mudskipper genus Oxuderces Eydoux & Souleyet, 1848 (Teleostei: Gobiidae: Oxudercinae) with resurrection from synonymy of O. nexipinnis (Cantor, 1849) – Zoological Journal of the Linnean Society, doi: 10.1111/zoj.12482

  • Jaafar Z., Polgar G., and Yuliadi Zamroni, 2016 – Description of a new species of Periophthalmus (Teleostei: Gobiidae) from the Lesser Sunda Islands – Raffles Bulletin of Zoology, 64: 278–283.

  • Jaafar Z., Perrig M. and Chou L.M., 2009 – Periophthalmus variabilis (Teleostei: Gobiidae: Oxudercinae), a valid species of mudskipper, and a re-diagnosis of Periophthalmus novemradiatusZoological Science, 26: 309–314.

  • Jaafar Z., and Larson H.L., 2008 – A new species of mudskipper, Periophthalmus takita (Teleostei: Gobiidae: Oxudercinae), from Australia, with a key to the genus – Zoological Science, 25: 946–952.

  • Jaafar Z., Lim K.K.P. and Chou L.M., 2006 – Taxonomical and morphological notes on two species of mudskippers, Periophthalmus walailakae and Periophthalmodon schlosseri (Teleostei: Gobiidae) from Singapore – Zoological Science, 23: 1043–1047.

  • Jayne B.C., Voris H.K., Heang K.B., 1988 – Diet, feeding behavior, growth and numbers of a population of Cerberus rynchops (Serpentes: Homalopsinae) in Malaysia – Fieldiana, 50: 1–15.

  • Jayne B.C., Ward T.J., Voris H.K., 1995 – Morphology, reproduction and diet of the marine homalopsine snake Bitia hydroides in Peninsula Malaysia – Copeia, 1995: 800–808.

  • Johnstone J., 1903 – Report on the Marine Fishes – In: Annadale N. and Robinson H. (eds) – Fasciculi Malayenses, Zoology – Part II. Longmans, Green & Company, New York.

  • Jordan D. and Seale A., 1906 – The fishes of Samoa – Bulletin of the U.S. Bureau of Fisheries – 25: 175–455.




  • Karsten, H., 1923 – Das Auge von Periophthalmus koelreuteriJenaische Zeitschrift für Naturwissenschaft, 59: 115–154.

  • Khaironizam, M.Z. and Norma–Rashid, Y., 2000 – A new record of the mudskipper Parapocryptes serperaster (Oxudercinae: Gobiidae) from Peninsular Malaysia – Malaysian Journal of Science, 19: 101–104.

  • Khaironizam, M.Z. and Norma–Rashid, Y., 2003 – First record of the mudskipper, Periophthalmodon septemradiatus (Hamilton) (Teleostei: Gobiidae) from Peninsular Malaysia – Raffles Bulletin of Zoology, 51(1): 97–100.

  • Khaironizam, M.Z. and Norma–Rashid, Y., 2002a – A new record of mudskipper Periophthalmus spilotus (Gobiidae: Oxudercinae) from Selangor coast, Peninsular Malaysia – Natural History Bulletin of the Siam Society, 50(2): 245–250.

  • Khaironizam, M.Z. and Norma–Rashid, Y., 2002b – Diet of mudskippers of the genus Periophthalmus from the Klang Strait, Malaysia – Phang Siew Moi, Chong Ving Ching, Ho Sinn Chye, Noraieni Mokhtar, Jillian Ooi Lean Sim (eds) – Marine Science into the New Millennium: New Perspectives & Challenges – Asia–Pacific Conference on Marine Science and Technology – 12–16 May 2002, Kuala Lumpur: 223–228.

  • Khaironizam M.Z. and Norma–Rashid Y., 2002c – Length-weight relationship of mudskippers (Gobiidae: Oxudercinae) in the coastal areas of Selangor, Malaysia – Naga, WorldFish Center Quarterly, 25(3,4): 20–22.

  • Khoo, K.G., 1966 – Studies on the biology of periophthalmid fishes in Singapore – BSc thesis, University of Singapore, 61 pp.

  • Kim I.–S., 1997 – Illustrated encyclopedia of fauna and flora of Korea. Vol. 37. Freshwater fishes – Ministry of Education: 1–629.

  • King, R.P. and Udo, M.T., 1997a – Some aspects of the reproductive biology of the endangered mudskipper, Periophthalmus barbarus (Gobiidae) in Imo river estuary, Nigeria – Transactions of the Nigerian Society of Biology and Conservation, 5: 50–58.

  • King, R.P. and Udo, M.T., 1997b – Vegetational succession–mediated spatial heterogeneity in the environmental biology of Periophthalmus barbarus (Gobiidae) in the estuarine swamps of Imo river, Nigeria – International Journal of Surface Mining, Reclamation and Environment, 11: 151–154.

  • King, R.P. and Udo, M.T., 1998 – Seasonality in diet and foraging performance of the mudskipper, Periophthalmus barbarus (Gobiidae) in Imo river estuary, Nigeria – In: R.P. King and B.S. Moses (eds) – Fish and Fisheries of Southeastern Nigeria – Publications Committee, Department of Zoology, University of Uyo, Uyo, Nigeria: June, 1998: 66–75.

  • Kobayashi T., Dotsu, Y. and Takita, T., 1971 – Nest and Nesting Behaviour of the Mudskipper Periophthalmus cantonensis in the Ariake Sound – Bulletin of the Faculty of Fisheries, Nagasaki University, 32: 27–39.

  • Kobayashi T., Dotsu Y. and Miura N., 1972 – Egg development and rearing experiments of the larvae of the mud skipper, Periophthalmus cantonensis (in Japanese with English summary) – Bulletin of the Faculty of Fisheries, Nagasaki University, 33: 49–62.

  • Koelreuter, I.T., 1763 – Piscium rariorum e Museo Petropolitano exceptorum descriptiones. Novi Commentarii Academiae Scientiarum Imperialis Petropolitanae v. 8 (1760-1761): 404-430, Pl. 1-6. [according to the Catalog of Fishes by Eschmeyer, I.T. Koelreuter is the same person as J.G. Koelreuter.]

  • Kok W.K., Lim C.B., Lam T.J. and Ip Y.K., 1998 – The mudskipper Periophthalmodon schlosseri respires more efficiently on land than in water and vice versa for Boleophthalmus boddaerti J. Exp. Zool., 280: 86–90.

  • Kottelat M., 2013 – The fishes of the inland waters of southeast Asia: a catalogue and core bibiography of the fishes known to occur in freshwaters, mangroves and estuaries – Raffles Bulletin of Zoology, Supplement No. 27: 1-663.

  • Kottelat M., Whitten A.J., Kartikasari S.N. and Wirjoatmodjo S., 1993 – Freshwater fishes of Western Indonesia and Sulawesi – Periplus Editions, Hong Kong. 221 p.

  • Kotrschal K., 1999 – Sensory Systems – In: Horn M.H., Martin K.L.M. and Chotkowski M.A. (eds) – Intertidal Fishes: Life in Two Worlds – Academic Press, San Diego California, 399 pp: 126–142.

  • Koumans F.P., 1937– Notes on Gobioid fishes 10. On a collection of the Museum of Basle, with description of a new species of ApocryptodonZoologisches Mededeelingen, Leiden Museum, 20(1938): 24–26.

  • Koumans F.P., 1941 – Gobioid fishes of India – Memoirs of the Indian Museum – v. 13 (pt 3): 205–329.

  • Koumans F.P., 1953 – Gobioidea – In: M. Weber and L. de Beaufort (eds.) – Fishes of the Indo–Australian Archipelago Vol. 10 – E.J. Brill, Leiden, 423 pp.

  • Koumans F.P., 1955 – Gobioid fishes of India – Memoirs of the Indian Museum, 13(1938–1942):205–330.

  • Kuang, D.H. and Y. Ni, 1986 – The freshwater and estuaries fishes of Hainan Island – Guangdong, China: Guangdong Science and Technology Press. 1-13, 1-3.




  • Larson H.K. and Lim K., 2004 (eds) – A Guide to Gobies of Singapore – Omni–Theatre, Singapore Science Centre, Singapore: 164 pp.

  • Larson H.K., 2008 – Weber's Mudskipper Periophthalmus weberi: New record for the Daly River – Northern Territory Naturalist, 20: 19–21.

  • Larson H.K., Jaafar Z. and Lim K.K.P., 2008 – An annotated checklist of the gobioid fishes of Singapore – The Raffles Bulletin of Zoology, 56(1): 135–155.

  • Larson H.K. and Takita T., 2004– Two new species of Periophthalmus (Teleostei: Gobiidae: Oxudercinae) from northern Australia, and a re–diagnosis of Periophthalmus novaeguineaensisThe Beagle, Records of the Museums and Art Galleries of the Northern Territory, 20: 175–185.

  • Lee Y.–J., Choi Y. and Ryu B.–S., 1995 – A taxonomic revision of the genus Periophthalmus from Korea with a description of a new species – The Korean Journal of Ichthyology, 7 (2): 120–127.

  • Lee H.J. and Graham B.J., 2002 – Their Game is Mud – Natural History, 9/02: 42–47.

  • Lee H.J., Martinez C.A., Hertzberg K.J., Hamilton A.L. and Graham J.B., 2005 – Burrow air phase maintenance and respiration by the mudskipper Scartelaos histophorus (Gobiidae: Oxudercinae) – The Journal of Experimental Biology, 208: 169–177.

  • Lesson R. P., 1830–1831 – Poissons – In: L. I. Duperrey – Voyage autour du monde, ..., sur la corvette de La Majesté La Coquille, pendant les années 1822, 1823, 1824 et 1825..., Zoologie –. Voyage Coquille Zool. v. 2 (pt 1): 66–238, Atlas: Pls. 1–38.

  • Leviton A.E., Gibbs R.H. Jr, Heal E. and Dawson C.E., 1985 – Standards in herpetology and ichthyology. Part 1. Standard symbolic codes for institutional resource collections in herpetology and ichthyology – Copeia, 3: 802–832.

  • Lim L.S., 1971 – The feeding biology of Periophthalmus vulgaris (Eggert) and Periophthalmodon schlosseri (Pallas) Perciformes, Periophthalmidae – MSc thesis, University of Malaya, 33 pp.

  • Lim C.B., Chew S.F., Anderson P.M. and Ip Y.K., 2001 – Reduction in the rates of protein and aminoacid catabolism to slow down the accumulation of endogenous ammonia: a strategy potentially adopted by mudskippers (Periophthalmodon schlosseri and Boleophthalmus boddarti) during aerial exposure in constant darkness – Journal of Experimental Biology, 204: 1605–1614.

  • Linnaeus C., 1758 – Systema naturae per regna tria naturae, secundum Classes, Ordines, Genera, Species, cum characteribus, differentiis, synonymis, locis (Pisces) – Laurentii Salvii, Holmiae (edn 10): 824 pp.

  • Linnaeus C., 1759 – Amoenitates academicæ seu dissertationes variæ physicæ, medicæ, botanicæ antehac seorsim editæ nunc collectæ et auctæ cum tabulis æneis – Laurentii Salvii, Holmiae (edn 12) Tomus 4, LXI. Chinensia Lagerström: 230–260.

  • Linnaeus C., 1766 – Systema naturae per regna tria naturae, secundum Classes, Ordines, Genera, Species, cum characteribus, differentiis, synonymis, locis (Pisces. Thoracici. Gobius) – Laurentii Salvii, Holmiae (edn 12) Tomus I: 532 pp.

  • Logan G. and Longmore A., 2003 – Sediment organic matters and nutrients – In: OzEstuaries Coastal Indicator Knowledge and Information System I: Biophysical Indicators – Canberra: Geoscience Australia. World Wide Web electronic publication, http://www.ozcoasts.org.au/indicators/sediment_org_matter.jsp (visited 14 April 2010).

  • Long A.J., 1995 (ed) – The Rise of Fishes – The John Hopkins University Press, Baltimore and London: 223 pp.

  • Long J.A. and Gordon M.S., 2004 – The Greatest Step in Vertebrate History: A Paleobiological Review of the Fish–Tetrapod Transition Physiological and Biochemical Zoology, 77(5): 700–719.

  • Low W.P., Ip Y. K. and Lane D.J.W., 1990 – A comparative study of the gill and skin morphometries of three mudskippers: Periophthalmus chrysospilos, Boleophthalmus boddaerti and Periophthalmodon schlosseri. Correlations and interpretations – Zoological Science, 7 (1): 29–38.




  • Macintosh D.J., 1977 – Some responses of tropical mangrove fiddler crabs (Uca spp.) to high environmental temperatures – In: McLusky D.S. & Berry A.J. (eds) – Physiology and Behaviour of Marine Organisms – Proceedings of the 12th European Symposium on Marine Biology. Stirling, Scotland, Pergamon Press: 49–56.

  • Macintosh D.J., 1979 – Predation of fiddler crabs (Uca spp.) in estuarine mangroves – Biotrop Special Publication No.10, Mangrove and Estuarine Vegetation in South East Asia, edited by P.B.L. Srivastava, Serdang, Malaysia: 101–110.

  • Macleay W., 1878 – The fishes of Port Darwin – Proc. Linn. Soc. N.S.W.: 344–367.

  • Macnae W. and Kalk M, 1962 – The ecology of the mangrove swamps at Inhaca Island, Mocambique. Journal of Animal Ecology, 50: 19–34.

  • Macnae W., 1968 – A General Account of the Fauna and Flora of Mangrove Swamps and Forests in the Indo–West–Pacific Region – Adv. Mar. Biol., 6: 73–270.

  • Magnus D.B.E., 1981 – Bewegungsweisen, Nahrungserwerb und Fortpflanzungverhalten des Schlammspringers Periophthalmus kalolo (Freilandaufnahmen) [Typical movement, Feeding and Mating Behaviour of the Mudskipper Periophthalmus kalolo (Open Air Shots)] – Publikationen zu Wissenschaftlichen Filmen, Sektion Biologie, Serie 14(7): Film D 1282, Institut für den Wissenschaftlichen Film (IWF), Göttingen.

  • Mansuri A.P., Bhan S. and Mathai T., 1982 – Salinity tolerance and survival of mudskipper Boleophthalmus dentatus and Periophthalmus dipes of Saurashtra coast – Journal of Animal Morphology and Physiology, 29(1-2): 142–148.

  • Martin K.L.M., 1996 – An ecological gradient in air–breathing ability among marine cottid fishes – Physiol. Zool., 69(5): 1096–1113.

  • Martin K.L.M. and Bridges C.R., 1999 – Respiration in air and water – In: Horn M.H., Martin K.L.M. and Chotkowski M.A. (eds) – Intertidal Fishes: Life in Two Worlds – Academic Press, San Diego California, 399 pp: 54–78.

  • Matoba M. and Dotsu Y., 1977 – Prespawning behavior of the mud skipper Periophthalmus cantonensis in Ariake Sound – Bulletin of the Faculty of Fisheries, Nagasaki University, 43: 23–33.

  • Mazda Y., Wolanski E., King B., Sase A., Ohtsuka D. and Magi, M., 1997a – Drag force due to vegetation in mangrove swamps – Mangroves & Salt Marshes, 1(3): 193–199.

  • Mazda Y., Michimasa M., Motohiko K. and Phan Nguyen Hong., 1997b – Mangroves as a coastal protection from waves in the Tong King delta, Vietnam – Mangroves & Salt Marshes, 1(2): 127–135.

  • Mazlan A.G. and Rohaya M., 2008 – Size, growth and reproductive biology of the giant mudskipper, Periophthalmodon schlosseri (Pallas, 1770), in Malaysian waters – Journal of Applied Ichthyology, 24(3): 290–296.

  • McCulloch A.R. and Waite E.R., 1918 – Descriptions of two new Australian Gobies – Records of the South Australian Museum, 1(1): 79–82.

  • Mhaisen F.T.; Al–Maliki N.S., 1996 – Parasites, diseases and food of the dark–blotched mudskipper Periophthalmus waltoni (Perciformes: Gobiidae) in the Khor Al–Zubair estuary (Iraq) – Zoology in the Middle East, 13: 85–87.

  • Milward N.E., 1974 – Studies on the taxonomy, ecology and physiology of Queensland mudskippers – unpub. Ph D. dissertation, Univ. Of Queensland, Brisbane, Australia, 276 pp.

  • Miller P.J., 1973 – The osteology and adaptive features of Rhyacicthys aspro (Teleostei:Gobioidei) and the classification of gobioid fishes – Journal of Zoology, London, 171: 397–434.

  • Miller P.J., 1984 – The Tokology of Gobioid Fishes – In: – Fish Reproduction: Strategies & Tactics – Academic Press Inc., Lond. Ltd: 119–153.

  • Miller P.J., 1984 – The Tokology of Gobioid Fishes – In: – Fish Reproduction: Strategies & Tactics – Academic Press Inc., Lond. Ltd: 119–153.

  • Miller P.J., 1986 – Reproductive Biology and Systematic Problems in Gobioid Fishes – In: – Indo–Pacific Fish Biology. Proceedings of the Second International Conference on Indo–Pacific Fishes, 1986 Ichthyological Society of Japan, Tokyo: 640–647.

  • Mirza M.R., Hameed S. and Naeem Javed M., 1996 – Fishes of Shadi Khoar (Baluchistan) – Second Symposium on Fish and Fisheries of Pakistan, November, 25, 26, 1996, Department of Zoology, Government College, Lahore, p. 8-9 (abstract).

  • Morii H. and Kasama K., 1989 – Bacterial Flora in the Digestive Tracts of Mudskipper Fishes Boleophthalmus pectinirostris and Periophthalmus cantonensisBulletin of the Japanese Society of Scientific Fisheries (Nippon Suisan Gakkaishi), 55(4): 733–741. (Japanese with English summary)

  • Morii H. and Kasama K., 1990 – The Character of Bacterial Flora in the Digestive Tract of Mudskipper Fishes Boleophthalmus pectinirostris and Periophthalmus cantonensisBulletin of the Faculty of Fisheries, Nagasaki University, 68: 43–52. (Japanese with English summary)

  • Munk O., 1970 – On the occurrence and significance of horizontal band–shaped retinal areae in teleosts – Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Khobenhavn, 133: 85–120.

  • Murdy E.O., 1986 – Mudskippers of Malaysia: The Lords of the Mudflat – Freshwater and Marine Aquarium Nov. 1986: 20–23.

  • Murdy E.O., 1989 – A Taxonomic Revision and Cladistic Analysis of the Oxudercine Gobies (Gobiidae: Oxudercinae) – Records of the Australian Museum Suppl. N°11: 1–93.

  • Murdy E.O., Takita T., 1999 – Periophthalmus spilotus, a new species of mudskipper from Sumatra (Gobiidae: Oxudercinae) – Ichthyological Research, 46 (4): 367–370.

  • Murphy D.C. (ed), 2005 – Devonian Times – World Wide Web electronic publication, www.devoniantimes.org, visited (11/2005).




  • Nanami A. and Takegaki T., 2005 – Age and growth of the mudskipper Boleophthalmus pectinirostris in Ariake Bay, Kyushu, Japan – Fisheries Research, 74: 24–34

  • Near T.J., Dornburg A., Eytan R.I., Keck B.P., Smith W.L., Kuhn K.L., Moore J.A., Price S.A., Burbrink F.T., Friedman M., Wainwright P.C., 2013 – Phylogeny and tempo of diversification in the superradiation of spiny-rayed fishes. &ndash Proceedings of the National Academy of Sciences USA, 110: 12738–12743.

  • Nelson J.S. (ed), 2006 – Fishes of the World – 4th edition. John Wiley. Hoboken.

  • Newman P.A., 2002 – An Introduction to Stratospheric Ozone (Cap. 1) – In: Todaro R.M. (ed), NASA's Goddard Space Flight Center Atmospheric Chemistry and Dynamics Branch (Code 916) http://www.nasa.gov/centers/goddard/home/index.htmlStratospheric Ozone – electronic book: http://www.ccpo.odu.edu/SEES/ozone/oz_class.htm visited (11/2005).

  • Nolf D. and Stringer G.L., 2003 – Late Eocene (Priabonian) Fish Otoliths from the Yazoo Clay at Copenhagen – Louisiana Geological Survey Geological Pamphlet No. 13: 1–23.

  • Nursall J.R., 1981 – Behaviour and Habitat Affecting the Distribution of Five Species of Sympatric Mudskippers in Queensland – Bulletin of Marine Science, 31 (3): 730–735.




  • Odum W.E. and Heald E.J., 1975 – The detritus–based food web of an estuarine mangrove community – In: Cronin G.E. (ed) – Estuarine Research, Vol. 1. Academic Press, New York: 265–286.

  • Odum E.P. (ed), 1983 – Basic Ecology – CBS College Publ.; Italian trad.: Basi di Ecologia, Piccin Nuova Libraria, Padova, 544 pp.

  • Oliva O. and Skorepa V., 1970a – The eye muscles in Periophthalmus Bloch & Schneider, 1801 and Chauliodus Bloch & Schneider, 1801 (Osteichthyes) – Vestnik Ceskoslovenske Spolecnosti Zoologicke Acta Societatis Zoologicae Bohemoslovacae, 34: 50–57.

  • Oliva O. and Skorepa V., 1970b – The eye muscles in Argyropelecus olfersi (Cuvier, 1829) and Boleophthalmus Val., 1837 – Vestnik Ceskoslovenske Spolecnosti Zoologicke Acta Societatis Zoologicae Bohemoslovacae, 34: 234–239.

  • Orton G.L., 1954 – Original adaptive significance of the tetrapod limb – Science, 120: 1042–1043.

  • Osbeck, P. 1765 – Reise nach Ostinden und China – xxvi + 578 pp, 13 pls.

  • Oshiro M., Tsuhako Y. and Tachihara K., 2005 – Habitat selection for burrow making by two species of mudskippers, Periophthalmus argentilineatus and P. modestus on the Okinawa Island, Japan – Okinawa Journal of Biology Education, 37: 9–18

  • Otto, A.W., 1821 – Conspectus animalium quorundam maritimorum, etc. Inaug. Dissert. Vratislaviae. Conspectus animalium quorundam maritimorum, etc.: 20 pp.




  • Pallas P.S., 1770 – Spicilegia Zoologica quibus novae imprimis et obscurae animalium species iconibus, descriptionibus atque commentariis illustrantur – Berolini, Gottl. August. Lange: 1–56.

  • Pan J.-H., Zhong L., Zheng C.-Y., Wu H.-L. and Liu J.-H. (eds) 1991 – The freshwater fishes of Guangdong Province – Guangdong Science and Technology Press. frontmatter + 1-589.

  • Pankow H. and Huq M.F., 1979 – Diatoms in the stomach content of Pseudapocryptes dentatus a mudskipper from the Shatt-al-Arab estuary (Iraq) – Wissenschaftliche Zeitschrift der Wilhelm-Pieck-Universität Rostock, Mathematisch-Naturwissenschaftliche, 28(6): 547–554.

  • Peng K.W., Chew S.F., Lim C.B., Kok T.W.K., Kuah S.S.L. and Ip Y.K., 1998 – The mudskippers Periophthalmus schlosseri and Boleophthalmus boddaerti can tolerate 446microM and 36microM of environmental NH3, respectively – Fish Physiology and Biochemistry, 19: 59–69.

  • (Gaius) Plinius Secundus, 77-79 AD – Naturalis Historia, Liber IX, XXXV.

  • Polgar G., 2004 – Adaptations to the peritidal environment of some Malaysian species of the genera Periophthalmus, Periophthalmodon and Boleophthalmus (Pisces, Gobiidae, Oxudercinae, Periophthalmini): ecological researches and realisation of a device for the rearing in semi–natural conditions – unpub. M.Sc. thesis (Laurea degree, summa cum laude), Univ. of Rome “La Sapienza”, Rome, Italy. (In Italian)

  • Polgar G., 2008 – Species-Area relationship and potential role as a biomonitor of mangrove communities of Malayan mudskippers – Wetlands Ecology and Management, DOI: 10.1007/s11273-008-9090-x (http://dx.doi.org/10.1007/s11273-008-9090-x)

  • Polgar G., 2016 – First record and conservation value of Periophthalmus malaccensis Eggert from Borneo, with ecological notes on other mudskippers (Teleostei: Gobiidae) in Brunei - Scientia Bruneiana, 15: 48-57 (paper online)

  • Polgar G. and Bartolino V., 2010 – Size variation of six species of oxudercine gobies along the intertidal zone in a Malayan coastal swamp – Marine Ecology and Progress Series, 409: 199-212 (DOI:10.3354/meps08597)

  • Polgar G., Burreson E.M., Stefani F. and Kamrani E., 2009 – Leeches on mudskippers: host-parasite interaction at the water's edge – Journal of Parasitology, 95(4): 1021-1025. (DOI: 10.1645/GE-1718.1)

  • Polgar G. and Crosa G., 2009 – Multivariate characterization of the habitats of seven species of Malayan mudskippers (Gobiidae: Oxudercinae) – Marine Biology, 156(7): 1475-1486. (DOI: 10.1007/s00227-009-1187-0)

  • Polgar G., Jaafar Z. and Konstantinidis P., 2013 – A new species of mudskipper, Boleophthalmus poti (Teleostei: Gobiidae: Oxudercinae) from the Gulf of Papua, Papua New Guinea, and a key to the genus – The Raffles Bulletin of Zoology: 61(1): 311-321 (http://rmbr.nus.edu.sg/rbz/biblio/61/61rbz311-321.pdf).

  • Polgar G. and Khaironizam M.Z., 2008 – First record of Periophthalmus walailakae (Gobiidae: Oxudercinae) from Peninsular Malaysia – Cybium, 32(4): 349–351.

  • Polgar G. and Lim R., 2011 – Mudskippers: human use, ecotoxicology and biomonitoring of mangrove and other soft bottom intertidal ecosystems – In: Metras J.N. (ed) Mangroves: Ecology, Biology and Taxonomy. Nova Science Publishers, Hauppauge, pp. 51-86. ISBN: 978-1-61728-991-0.

  • Polgar G., Malavasi S., Cipolato G., Georgalas V., Clack J.A. and Torricelli P., 2011 – Acoustic communication at the water's edge: Evolutionary insights from a mudskipper – PLoS ONE, 6(6): e21434. (doi:10.1371/journal.pone.0021434)

  • Polgar G., Sacchetti A. and Galli P., 2010 – Differentiation and adaptive radiation of amphibious gobies (Gobiidae: Oxudercinae) in semi-terrestrial habitats – Journal of Fish Biology, 77: 1645–1664 (DOI:10.1111/j.1095-8649.2010.02807.x)

  • Polgar G. and Sasekumar A., 2010 – The potential of ecotourism and flagship species, and the urgent need for more holistic approaches and higher levels of connectivity for endangered tropical mudflats and coastal swamps – In: Krause A. and Weir E. (eds) Ecotourism: management, development and impact. Nova Science Publishers, Hauppauge. ISBN: 978-1-60876-724-3.

  • Polgar G., Zane L., Babbucci M., Barbisan F., Patarnello T., Rüber L., and Papetti C. (2014) – Phylogeography and demographic history of two widespread Indo-Pacific mudskippers (Gobiidae: Periophthalmus) – Molecular Phylogenetics and Evolution, 73: 161-176.

  • Poll M., 1959 – Expédition océanographique belge dans les eaux côtières africaines de l'Atlantique Sud (1948-1949). Résultats scientifiques – Volume IV, fascicule 3B. Poissons. V. - Téléostéens acanthoptérygiens (deuxième partie). Institut Royal des Sciences Naturelles de Belgique, Bruxelles. 417 pp.

  • Polunin I. W., 1972 – Who Says Fish Cant' Climb Trees? – National Geographic, 141: 85–91.

  • Proches S., Polgar G. and Marshall D.J., 2014 – K-Pg events facilitated lineage transitions between terrestrial and aquatic ecosystems – Biology Letters, 10(6) doi: 10.1098/rsbl.2014.0010.




  • Quoy J.R.C. and Gaimard J.P., 1824–25 – Description des Poissons. Chapter IX – In: Freycinet, L. de – Voyage autour du Monde... exécuté sur les corvettes de L. M. "L'Uranie" et "La Physicienne," pendant les années 1817, 1818, 1819 et 1820 – Paris. Voyage Uranie, Zool.: 192–401 [1–328 in 1824; 329–616 in 1825], Atlas pls. 43–65.




  • Radke L.C., Smith C.S., Ryan D.A., Brooke B., Heggie D. and contributors, 2003 – Coastal Indicator Knowledge and Information System I: Biophysical Indicators – Canberra: Geoscience Australia. World Wide Web electronic publication, www.ozestuaries.org/indicators, visited (11/2005)

  • Rahman, A.K.A., 1989 – Freshwater fishes of Bangladesh – Zoological Society of Bangladesh. Department of Zoology, University of Dhaka: 364 pp.

  • Rainboth W.J., 1996 – Fishes of the Cambodian Mekong. – FAO Species Identification Field Guide for Fishery Purposes. FAO, Rome: 265 pp.

  • Randall D.J., Wilson J.M., Peng K.W., Kok T.W.K., Kuah S.S.L., Chew S.F., Lam T.J. and Ip Y.K., 1999 – The mudskipper, Periophthalmodon schlosseri, actively transports NH4+ against a concentration gradient – Am. J. Physiol. Regul. Integr. Comp. Physiol., 277(6): R1562–R1567.

  • Randall D.J. and Tsui T.K.N., 2002 – Ammonia toxicity in fish – Marine Pollution Bulletin, 45: 17–23.

  • Randall D.J., Ip Y.K., Chew S.F. and Wilson J.M., 2004 – Air Breathing and Ammonia Excretion in the Giant Mudskipper, Periophthalmodon schlosseri – Physiological and Biochemical Zoology, 77(5): 783–788.

  • Randall J.E., 1995 – Coastal fishes of Oman – University of Hawaii Press, Honolulu, Hawaii: 439 p.

  • Randall, J.E., 1997 – Randall's tank photos. Collection of 10,000 large–format photos (slides) of dead fishes. – Unpublished.

  • Raven P.H., Evert R.F. and Curtis H. (eds), 1981 – Biology of Plants – Worth Publishers Inc., New York; Italian edition 1984: Biologia delle piante, Nicola Zanichelli S.p.A., Bologna, 622 pp.

  • Ravi V., Rajagopal S., 2007 – Age and growth of the mudskipper, Bolephthalmus boddarti (Pallas, 1770). – Journal of Aquatic Biology, 22(1): 123–128.

  • Renema W., Bellwood D.R., Braga J.C., Bromfield K., Hall R., Johnson K.G., Lunt P., Meyer C.P., McMonagle L.B., Morley R.J., et al., 1980 – Hopping hotspots: global shifts in marine biodiversity – Science, 321: 654–657.

  • Ricci Lucchi F. and Mutti E. (eds), 1980 – Sedimentologia – Parti I–III, CLUEB, Bologna, 545 pp.

  • Richardson J., 1844–1845 – Ichthyology, Part 3 – In: R.B. Hinds (ed.) – The zoology of the voyage of H.M.S. Sulphur, under the command of Captain Sir Edward Belcher, R.N., C.B., F.R. G.S., etc., during the years 1836–42 – No. 10. London: Smith, Elder & Co.: 150 pp.

  • Richardson J., 1846 – Report on the ichthyology of the seas of China and Japan – Rep. Brit. Assoc. Adv. Sci.: 187–320.

  • Roberts T.R., 1978 – An ichthyological survey of the Fly River in Papua New Guinea with descriptions of new species – Smithson. Contrib. Zool., 281: 1–72.

  • Rochebrune A.T. (de), 1883 – Faune de la Sénégambie. Poissons – Actes de la Société de Bordeaux, 36(6) (1882): 20–190, Pls.

  • Romer A.S. and Parsons T.S. (eds), 1986 – The Vertebrate Body – Saunders College Publ., 6th ed.: 735 pp.




  • Sakamoto T., Yokota S and Ando M., 2000 – Rapid morphological oscillation of mitochondrion–rich cell in estuarine mudskipper following salinity changes – Journal of Experimental Zoology, 286: 666–669.

  • Sakamoto T. and Ando M., 2002 – Calcium ion triggers rapid morphological oscillation of chloride cells in the mudskipper, Periophthalmus modestusJournal of Comparative Physiology (B), 172: 435–439.

  • Salih M.S. and Al-Jaffery A., 1980 – Morphological study of Boleophthalmus dentatus Cuvier & Valenciennes, 1837 with reference to its feeding habits. 1: Habitat, external features and integument. Bulletin of the Biological Research Center, Baghdad, 12(2): 11–44.

  • Sarker A.L., Al-Daham N.K. and Bhatti M.N., 1980 – Food habits of the mudskipper, Pseudapocryptes dentatus (Val.). Journal of Fish Biology, 17(6): 635–639.

  • Sasekumar A., 1980 – The Present State of Mangrove Ecosystems in SouthEast Asia and the Impact of Pollution – FAO/UNEP project: Impact of Pollution on the Mangrove Ecosystem and its Productivity in Southeast Asia; (SCS/80/WP/94b). Publication of the South China Sea Fisheries Development and Coordinating Programme, 4–8 Feb. 1980, Manila, Philippines, 80 pp.

  • Sasekumar A., Ong T.L. and Thong K.L., 1984 – Predation of mangrove fauna by marine fishes – Soepadmo E., Rao, A.N. & Macintosh D.J. (eds). Proceedings of the Asean Symposium on Mangrove Environment. Research and Management. University of Malaya and UNESCO, Kuala Lumpur, Malaysia, pp. 378–384.

  • Sasekumar A., Chong V.C., Lim, K.H. and Singh H.R., 1994 – The Fish Community of Matang Mangrove Waters, Malaysia – Sudara, S.; Wilkinson, C.R.; Chou, L.M. (eds). Proceedings, Third ASEAN–Australia Symposium on Living Coastal Resources. Research papers. Chulalonghorn University, Bangkok, Thailand. Vol. 2: 457–464.

  • Saxon J., 1975 – The Fossil Fishes of the North of Scotland – John Humphries, Caithness Books, Thurso.

  • Schultze H.–P., 1999 – The Fossil Record of the Intertidal Zone – In: Horn M.H., Martin K.L.M. & Chotkowski M.A. (eds) – Intertidal Fishes: Life in Two Worlds – Academic Press, San Diego California, 399 pp: 373–392.

  • Schwab I.R., 2003 – Janus on the mudflats – British Journal of Ophthalmology, 87: 13.

  • Smith M.M. and Heemstra P.C., 1986 – Smiths' sea fishes – Springer–Verlag, Berlin.

  • Smith W.L. and Craig M.T., 2007 – Casting the percomorph net widely: the importance of broad taxonomic sampling in the search for the placement of serranid and percid fishes – Copeia, 2007: 35–55.

  • Smith W.L. and Wheeler W.C., 2004 – Polyphyly of the mail-cheeked fishes (Teleostei: Scorpaeniformes): evidence from mitochondrial and nuclear sequence data – Molecular Phylogenetics and Evolution, 32: 627–646.

  • Soni V.C. and George B., 1986 – Age determination and length-weight relationship in the mudskipper Boleophthalmus dentatus. Indian Journal of Fisheries, 33: 231–234.

  • Špinar Z.V. and Burian Z., 1972 – Life Before Man – Artia, Praga; Italian trad.: Quando l'uomo non c'era, Fratelli Fabbri Editori, Milano, 228 pp.

  • Sponder D.L. and Lauder G.V., 1981 – Terrestrial Feeding in the Mudskipper Periophthalmus (Pisces: Teleostei). A Cineradiographic Analysis – Journal of Zoology, Lond., 193 (4): 517–530.

  • Springer V.G., 1978 – Synonymization of the family Oxudercidae, with comments on the identity of Apocryptes cantoris Day (Pisces: Gobiidae) – Smithsonian Contributions to Zoology, 270: i-iii + 1-14.

  • Stebbins R.C. and Kalk M., 1961 – Observations on the Nat. Hist. of the Mud–skipper, Periophthalmus sobrinus Copeia, 1: 18–27.

  • Stille P., Steinmann M. and Riggs S.R., 1996 – Nd isotope evidence for the evolution of the paleocurrents in the Atlantic anc Tethys Oceans during the past 180 Ma – Journal of Experimental Zoology, 284 (3): 265–270.

  • Swainson W., 1839 – Natural History and Classification of Fishes, Amphibians and Reptiles – Vol. II. Longman, Orme, Brown, Green, & Longmans and John Taylor, London: 452 pp.

  • Swennen C. Ruttanadakul N., Haver M., Piummongkol S, Prasertsongscum S., Intanai I., Chaipakdi W., Yeesin P., Horpet P. and Detsathit S., 1995 – The Five Sympatric Mudskippers (Teleostei: Gobioidea) of Pattani area, Southern Thailand – Nat. Hist. Bull. Siam. Soc., 42: 109–129.

  • Swofford D.L., 2002 – Phylogenetic Analysis Using Parsimony, version 40b10 – Sunderland, Sinauer Assoc.




  • Takeda T., Ishimatsu A., Oikawa S., Kanda T., Ishida Y. and Khoo K.H., 1999 – Mudskipper Periophthalmodon schlosseri can repay oxygen debts in air but not in water – Earth and Planetary Science Letters , 144(1–2): 9–19.

  • Takegaki, T., Fujii T. and Ishimatsu A., 2006 – Overwintering habitat and low-temperature tolerance of the young mudskipper Boleophthalmus pectinirostrisNippon Suisan Gakkaishi, 72: 880–885 (In Japanese with English abstract).

  • Takegaki, T., 2008 – Threatened fishes of the world: Boleophthalmus pectinirostris (Linnaeus 1758) (Gobiidae) – Environmental Biology of Fishes, 81(4): 373–374.

  • Takita T., Agusnimar and Ahyudin B. Ali, 1999 – Distribution and habitat requirements of oxudercine gobies (Gobiidae: Oxudercinae) along the Straits of Malacca – Ichthyological Research, 46 (2): 131–138.

  • Takita T., Larson H.K. and Ishimatsu A., 2011 – The natural history of mudskippers in northern Australia, with field identification characters – The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 27: 189–204.

  • Talwar P.K. and Jhingran A.G., 1991 (eds) – Inland fishes of India and adjacent countries – Volume 2. A.A. Balkema, Rotterdam, pp. 962–963.

  • Taylor J.R., Cook M.M., Kirkpatrick A.L., Galleher S.N., Eme J. and Bennett W.A., 2005 – Thermal Tactics of Air–Breathing and Non Air–Breathing Gobiids Inhabiting Mangrove Tidepools on Pulau Hoga, Indonesia – Copeia, (4): 886–893.

  • Thacker C.E., 2003 – Molecular phylogeny of the gobioid fi shes (Teleostei: Perciformes: Gobioidei) – Molecular Phylogenetics and Evolution, 26: 354–368.

  • Thacker C.E., Satoh T.P., Katayama E., Harrington R.C., Eytan R.I., Near T.J., 2003 – Molecular phylogeny of Percomorpha resolves Trichonotus as the sister lineage to Gobioidei (Teleostei: Gobiiformes) and confirms the polyphyly of Trachinoidei – Molecular Phylogenetics and Evolution, 93: 172–179.

  • Tomiyama I., 1934 – Four new species of gobies of Japan – J. Fac. Sci. Univ. Tokyo, Sect. IV Zool.: 325–334.

  • Tomiyama I. and Abe T., 1958 – Pseudapocryptes lanceolatus (Bloch & Schneider). In: Tomiyama I. and Abe T. (eds) Fishes and descriptions of the Fishes of Japan (a continuation of Dr. Shigeo Tanaka's work), Vol LIX, CCCXXXV, p. 1231-1235; plate CCXXXVII – Shobo K. Publ. Tokyo, Japan.

  • Tornabene L., Chen Y., Pezold F., 2013 – Gobies are deeply divided: phylogenetic evidence from nuclear DNA (Teleostei: Gobioidei: Gobiidae) – Systematics and Biodiversity, 11(3): 345–361.

  • Townsend K.A. and Tibbetts I.R., 1995 – Re–appearance of the blue mudhopper, Scartelaos histophorus in the Greater Brisbane area – Memoirs of the Queensland Museum, 38 (2): 671–676.

  • Townsend K.A. and Tibbetts I.R., 2005 – Behaviour and sexual dimorphism of the blue mudskipper, Scartelaos histophorus (Pisces: Gobiidae) – Proceedings of the Royal Society of Queensland, 112: 53 –62.

  • Tsuhako Y., Ishimatsu A., Takeda T., Khoo K.H. and Tachihara K., 2003 – The eggs and larvae of the giant mudskipper, Periophthalmodon schlosseri, collected from a mudflat in Penang, Malaysia – Ichthyological Research, 50: 178–181.

  • Tytler P. and Vaughan T., 1983 – Thermal Ecology of the Mudskippers Periophthalmus koelreuteri (Pallas) and Boleophthalmus boddaerti (Pallas), of Kuwait Bay – Journal of Fish Biology, 23(3): 327–337.




  • UNEP–WCMC Coral Reefs and Mangroves of the World, 2005 – World distribution of coral reefs and mangroves – World Wide Web electronic publication, www.unep–wcmc.org/marine/data/coral_mangrove/index.html, visited (10/2005).

  • Ultsch, G.R., 1996 – Gas exchange, hypercarbia and acid–base balance, palaeoecology and the evolutionary transition from water breathing to air–breathing among vertebrates – Palaeogeography, Palaeoclimatology, Palaeoecology, 123: 1–27.




  • Wang H.Y., Tsai M.–P., Dean J. and Lee S.C., 2001 – Molecular Phylogeny of Gobioid Fishes (Perciformes: Gobioidei) Based on Mitochondrial 12S rRNA Sequences – Molecular Phylogenetics and Evolution, 20 (3): 390–408.

  • Washio M., Tsutsui M. and Takita T., 1991 – Age and growth of the mudskipper Boleophthalmus pectinirostris distributed in the mud flat of the Midori river, Kumamoto prefecture – Bulletin of the Japanese Society of Scientific Fisheries (Nippon Suisan Gakkaishi), 57(4): 637–644. (In Japanese with English summary)

  • Weber M. and De Beaufort L.F., 1953 – The fishes of the Indo–Australian archipelago. X. Gobioidea – A.J. Reprints Agency, New Delhi, 423 pp.

  • Winterbottom R., 1993 – Search for the gobioid sister group (Actinopterygii: Percomorpha) – Bulletin of Marine Science, 52 (1): 395–414.

  • White T., Clowes C., Haaramo M., Kazlev M.A. and Taylor C. (eds), 2005 – Palaeos: The Trace of Life on Earth – World Wide Web electronic publication, palaeos.com, visited (11/2005).

  • Whitley G.P. 1931 – New names for Australian fishes – Australian Zoologist, 6: 310–334.

  • Whitley G.P. 1953 – Fishes collected by the Australian Museum Expedition, 1952 – Records of the Australian Museum, 19: 60–112.

  • Whitley G.P. 1961 – A new mud–skipper from Papua (Pisces: Periophthalmidae) – Proc. R. Zool. Soc. N. S. W. , v. for 1958–59: 69–70.

  • Whittaker R.H. and Likens G., 1973 – Carbon in the biota – In: Woodwell G. & Pecan E. (eds) – Carbon and the Biosphere – CONF 720510, National Technical Information Service, Washington, D.C.

  • Wu H.–W., 1931 – Notes on the fishes from the coast of Foochow Region and Ming River – Contributions from the Biological Society of China, Zoological Series, 7(1): 1–64, 10 figs.




  • Yang, K.Y.J., 1996 – The feeding ecology of the mudskipper Boleophthalmus pectinirostris (Pisces: Periophthalmidae) at the Mai Po marshes nature reserve, Hong Kong – Thesis for the degree of Master of Phylosophy. Faculty of Science, Hong Kong University. 193 pp.

  • Yang K.Y.; Lee, S.Y. and Williams G.A., 2003 – Selective feeding by the mudskipper (Boleophthalmus pectinirostris) on the microalgal assemblage of a tropical mudflat – Marine Biology, 143: 245–256.

  • Yokota S., Iwata K., Fujii Y. and Ando M., 1997 – Ion transport across the skin of the mudskipper Periophthalmus modestus – Comp. Biochem. Physiol. 118A, 903–910.




  • Zander C.D., Nieder J. and Martin K., 1999 – Vertical Distribution Patterns – In: Horn M.H., Martin K.L.M. & Chotkowski M.A. (eds) – Intertidal Fishes: Life in Two Worlds – Academic Press, San Diego California, 399 pp: 26–53.

  • Zhang H.–G. and He D., 1989 – [The comparison of the retino–motor response of Epinephelus fario and Boleophthalmus pectinirostris under various light conditions] – Journal of Xiamen University, Natural Science, 28: 647–650. (In Chinese with English abstract)

  • Zhang Q.–Y., Hong W.–S., Dai Q.–N., Cai Y.–Y. and Zhang J., 1987 – Studies on the Artificial Propagation and Larval Rearing of the Mudskipper Boleophthalmus pectinirostris L. – J. Xiamen Univ. (Nat. Sci.) /(Xiamen Daxue Xuebao), 26(3): 366–373. (In Chinese with English summary)

  • Zhang Q.–Y., Hong W.–S., Dai Q.–N., Zhang J., Cai Y.–Y. and Huang J.–L., 1989 – Studies on induced ovulation, embryonic development and larval rearing of the mudskipper (Boleophthalmus pectinirostris) – Aquaculture, 83: 375–385.

  • Zhang Q.–Y. and Zhang J., 1988 – On the Feeding, Growth and Survival of Larval Mudskippers (Boleophthalmus pectinirostris) – J. Fish. China (Shuichan Xuebao), 12(3): 203–212. (In Chinese with English summary)

  • Zhang J., Taniguchi T., Takita T., Ahyaudin B. Ali, 2000 – On the epidermal structure of Boleophthalmus and Scartelaos mudskippers with reference to their adaptation to terrestrial life – Ichthyological Research, 47(3-4): 359–366.

  • Zhang J., Taniguchi T., Takita T. & Ahyaudin B. Ali, 2003 – A study on the epidermal structure of Periophthalmodon and Periophthalmus mudskippers with reference to their terrestrial adaptation – Ichthyological Research, 50: 310–317.

  • Zhu M. and Ahlberg P.E., 2004 – The origin of the internal nostril of tetrapods– Nature, 432: 94–97.




Creative Commons License
All content on this website (including text, photographs, and any other original works), unless otherwise noted, is licensed under a Creative Commons License